Drainage reorganization in intracontinental basins has strong implications on the sediment routing system and on the landscape evolution at a cratonic scale. Fluvial captures have strong impact on drainage areas, fluxes, and so on incision capacity, especially for the captured basin. Although the two Ebro and Duero hydrographic basins recorded similar histories, they are characterized by very different morphologic features. The Ebro basin is highly excavated, whereas the Duero basin is well preserved and may be considered as almost still endorheic. These two bordering basins then show contrasting preservation states of their endorheic stages and represent an ideal natural laboratory to study what factors (internal / external) control drainage divide mobility, and drainage network and landscape evolution in post-orogenic basins.